The editors of Zoological Research gratefully acknowledge the generous assistance of the following reviewers. We are thankful for the honest and invaluable work of all the referees, which has greatly contributed to ensuring the quality of Zoological Research.
Hakan Gur
Ahi Evran University, Turkey
Masakazu Asahara
Aichi Gakuin University, Japan
Gul Olgun Karacan
Aksaray University, Turkey
Russell J. Ferland
Albany Medical College, USA
Alex Matsyura
Altai State University, Russia
Saeed Seifi
Amol University of Special Modern Technologies, Iran
Hong-Chun Pan
Anhui Normal University, China
Jin-Hua Li Bao-Wei Zhang Dong-Po Xia
Anhui University, China
Krishnanand Nagarajan
Anna University of Technology, India
Xu Wang
Auburn University, USA
Donald J.Colgan
Australian Museum, Australia
Aurelio Ramírez-Bautista
Autonomous University of Hidalgo, Mexico
Jeffrey Rogers
Baylor College of Medicine, USA
Wei-Dong Bao Ji-Liang Xu
Beijing Forestry University, China
Yong-Liang Zhao
Beijing Institute of Genomics, Chinese Academy of Sciences, China
Ding-Zhen Liu Zheng-Wang Zhang
Beijing Normal University, China
Georgy Shenbrot
Ben-Gurion University of the Negev, Israel
Shih-Pin Huang
Biodiversity Research Center, Academia Sinica, Nankang, Taipei, Taiwan, China
Seth M. Bybee
Brigham Young University, USA
Michael B. Harvey
Broward College, USA
David Glenn Smith
California National Primate Research Center, USA
Cynthia M. Beall
Case Western Reserve University, USA
Hai-Peng Li
CAS-MPG Partner Institute for Computational Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, China
Hua Wu
Central China Normal University, China
Mrinal Samanta
Central Institute of Freshwater Aquaculture, India
Zuo-Fu Xiang
Central South University of Forestry and Technology, China
Lori Sheeran
Central Washington University, USA
Jean-Philip Brugal
Centre National de la Recherche Scientifique, France
Luca Luiselli
Centre of Environmental Studies Demetra, Italy
Daniel Frynta
Charles University, Czech Republic
Jia-Tang Li Bin Wang Yu-Chi Zheng
Chengdu Institute of Biology, Chinese Academy of Sciences, China
Jian-Yong Han
China Agricultural University, China
Hai Qian
China Pharmaceutical University, China
Xiaoyu Tian
Chinese University of Hong Kong, China
Da-Yong Li Wei Wei
China West Normal University, China
Christina Wilkinson
Clemson University, USA
Shahrina Chowdhury
City University of New York, USA
Noah T. Dunham
Cleveland Metroparks Zoo, USA
Ma. Teresa Santillan-Galicia
Colegio de Postgraduados, Mexico
Lei Yang
College of Charleston, USA
Thomas Ziegler
Cologne Zoo, Germany
Agustín G. Zapata
Complutense University, Spain
Sarah Sanders Lower Yun-Ke Wu
Cornell University, USA
Gerhard F. Weinbauer
Covance Preclinical Services, Germa
Wen Xiao
Dali University, China
Julie Iskander
Deakin University, Australia
Jitendra Sharma
Dibrugarh University, India
Yang-Zhong Huang George Tiley
Duke University, USA
Xiu-Guang Mao
East China Normal University, China
Georgette Gafford
Emory University School of Medicine, USA
Pascal Fries
Ernst Strüngmann Institute (ESI) for Neuroscience in Cooperation with Max Planck Society, Germany
Julian Kerbis
Field Museum of Natural History, USA
Robert W. Stackman
Florida Atlantic University, USA
Manxiu Ma
Fralin Biomedical Research Institute at Virginia Tech Carilion, Virginia Tech, USA
Sebastian Guenther
Free University Berlin, Germany
Jiu-Cun Wang Biao Yan
Fudan University, China
Stephen Greiman
Georgia Southern University, USA
David A. Washburn
Georgia State University, USA
Moein Esghaei
German Primate Centre, Leibniz Institute for Primate Research, Germany
Li-Biao Zhang
Guangdong Institute of Applied Biological Resources, China
Qi-Hai Zhou
Guangxi Normal University, China
Wei-Cai Chen
Guangxi Teachers Education University, China
Jiang Zhou
Guizhou Normal University, China
Wei Liang
Hainan Normal University, China
Wim Vanduffel
Harvard Medical School, USA
David Haig
Harvard University, USA
Ulrich Lehmann
Hannover Medical School, Germany
Bao-Liang Fan
Hebei Agricultural University, China
Feng Zhang
Hebei University, China
Liora Kolska Horwitz
Hebrew University of Jerusalem, Israel
Yoram Ben-Shaul
Hebrew University of Jerusalem Faculty of Medicine, Israel
Jian-Li Xiong
Henan University of Science and Technology, China
Hitoshi Suzuki
Hokkaido University, Japan
Lejla Daruka
Hungarian Academy of Science Biological Research Centre, Hungary
David Tarkhnishvili
Ilia State University, USA
Deepak Veerappan
Indian Institute of Science, India
Kunjbihari Sulakhiya
Indira Gandhi National Tribal University, Amarkantak, India
Amir Hamidy
Indonesian Institute of Sciences, Indonesia
Yong-E Feng
Inner Mongolia Agricultural University, China
Jérome Casas
Institut de Recherche sur la Biologie de l'Insecte, France
Frederic Ariey
Institut Pasteur, France
Olivier S. G. Pauwels
Institut Royal des Sciences Naturelles de Belgique, Belgium
Yuri Marusik
Institute for Biological Problems of the North, Russian Academy of Sciences, Russia
Bing Liu
Institute of Automation, Chinese Academy of SciencesChina
Ning Liu
Institute of Biophysics, Chinese Academy of Sciences, China
Gabriele Schino
Institute of Cognitive Sciences and Technologies, Italy
Truong Quang Nguyen
Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, Vietnam
Wen-Feng Qian
Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, China
Shun-Ping He Huan-Zhang Liu Pin Nie E Zhang
Institute of Hydrobiology, Chinese Academy of Sciences, China
Wei Cun Jie-Jie Dai Cun-Bao Liu Long-Ding Liu
Institute of Medical Biology, Chinese Academy of Medical Science and Peking Union Medical College, China
Song-Tao Yang
Institute of Military Veterinary, Academy of Military Medical Sciences, China
Yong Gu
Institute of Neuroscience, Chinese Academy of Sciences, China
Zhong-Li Sha
Institute of Oceanology, Chinese Academy of Sciences, China
Patrice Brehmer
Institute of Research for Development, France
Sang Nguyen
Institute of Tropical Biology, Vietnam Academy of Science and Technology, Vietnam
De-Yan Ge Yun-Biao Hu Cheng-Ming Huang Zhi-Gang Jiang Lei Li Ming Li Bao-Ping Ren Yan Xie De-Hua Wang Jian-Guo Zhao Ya-Hui Zhao
Institute of Zoology, Chinese Academy of Sciences, China
Juan J. Calvete
Instituto de Biomedicina de Valencia, Spain
Juan C. Cancino-Diaz
Instituto Politécnico Nacional, Mexico
Russell A. Kern
International Stem Cell Corporation, USA
Rasoul Kowsar
Isfahan University of Technology, Iran
Qing-Pi Yan
Jimei University, China
Wan-Sheng Jiang Zhi-Xiao Liu
Jishou University, China
Jian-Huan Yang
Kadoorie Farm and Botanic Garden, Hong Kong, China
PingWu
Keck Schoolof Medicine of the University of Southern California, USA
Niti Sukumal
King Mongkut's University of Technology Thonburi, Thailand
Yoji Osako
Kochi Medical School, Japan
Yu-Peng Cun Hua-Ying Sun
Kunming Institute of Botany, Chinese Academy of Sciences, China
Ce-Shi Chen Xiao-Yong Chen Li-Na Du Kai He Ke Jiang Xue-Long Jiang Bao-Wei Jiao Xue-Yan Li Bin Liang Xiong-Jian Luo Bing-Yu Mao Min-Sheng Peng Neng-Yin Sheng Yong-Gang Yao Dong-Dong WU Guo-Jie Zhang Yun Zhang Zhi-Ye Zhang
Kunming Institute of Zoology, Chinese Academy of Sciences, China
Yong-Chang Chen
Kunming University of Science and Technology, China
Sannanegunda V. Krishnamurthy
Kuvempu University, India
Koshiro Eto Takuro Ikeda Masaharu Motokawa Kanto Nishikawa
Kyoto University, Japan
Takeshita Rafaela
Kyoto University Primate Research Institute, Japan
Jin-Won Lee
Kyung Hee University, Korea
Hiroyuki Sasaki
Kyushu University, Japan
Grismer L. Lee
La Sierra University, USA
Qiang Qiu
Lanzhou University, China
Fabian Herder
Leibniz Institute for Animal Biodiversity, Germany
Renata F. Martins
Leibniz Institute for Zoo and Wildlife Research, Germany
Klaus Kohlmann
Leibniz Institute of Freshwater Ecology and Inland Fisheries, Germany
Michael K. Richardson
Leiden University, Netherlands
Nikolay A. Poyarkov, Jr. Eduard Galoyan
Lomonosov Moscow State University, Russia
Suresh Balakrishnan
Maharaja Sayajirao University of Baroda, India
Achom Darshan
Manipur University, India
James J. DiCarlo
Massachusetts Institute of Technology, USA
Achyut Aryal
Massey University, New Zealand
Anneke Janzen
Max-Planck-Institut fur Menschheitsgeschichte, Germany
Jean-Pierre Routy Elena Torban Kim Valenta
McGill University, Canada
Iwona Zarnowska
Medical University, Poland
Gonzalo J. Revuelta Wei Jiang
Medical University of South Carolina, USA
Malinalli Cortes-Marcial
Metropolitan Autonomous University, Mexico
Xin Du
Monash University, Australia
Fabio Machado
Museo Argentino de Ciencias Naturales Bernardino Rivadavia, Argentina
Ana L.C. Prudente
Museu Paraense Emílio Goeldi, Brazil
Helen Larson
Museum and Art Gallery of the Northern Territory, Australia
Annemarie Ohler David Patrick Gael P. J. Denys
Museum National d'Histoire Naturelle, France
Mya Myat Ngwe Tun
Nagasaki University, Japan
Takuma Kaito
Naha Nature Conservation Office, Ministry of the Environment, Japan
Xiao-Ping Wu
Nanchang University, China
Xiang JI
Nanjing Normal University, China
Xiang Gao
Nanjing University, China
Jian Yang
Nanning Nornal University
Joseph Koh
National Biodiversity Centre, Singapore
Takaaki Suzuki
National Institute for Environmental Studies, Japan
William J. Liu
National Institute for Viral Disease Control and Prevention,Chinese Center for Disease Control and Prevention, China
Hidetoshi Inagaki Yoshihiro Ohmiya
National Institute of Advanced Industrial Science and Technology (AIST), Japan
Junghwa An
National Institute of Biological Resources, Korea
Masayuki Saijo
National Institute of Infectious Diseases, Japan
Laxmi T. Rao
National Institute of Mental Health & Neuro Science, India
Monalisa Mishra
National Institute of Technology Rourkela, India
Esther Kioko
National Museums of Kenya, Kenya
Hiroshi Morino
National Museum of Nature and Science, Japan
John D. Harding
National Primate Research Centers, USA
Judie Alimonti
National Research Council of Canada, Canada
Jhan-Wei Lin
National Taiwan Normal University, China
Keng-Chen Liang
National Taiwan University, China
Cedric Finet Heok Hee Ng Le Wang
National University of Singapore, Singapore
Eric Nelson
Nationwide Children’s Hospital, USA
Ignasi Torre
Natural History Museum of Granollers, Spain
Gareth Richards
Newcastle University, UK
Hirohiko Takeuchi
Nihon University, Japan
Jiong Chen
Ningbo University, China
Bryan L. Stuart
North Carolina State Museum of Natural Sciences, USA
Craig V. Sullivan
North Carolina State University, USA
Erin Gillam
North Dakota State University, USA
Yu Jiang Chu-Zhao Lei
Northwest A & F University, China
Song-Tao Guo Bao-Guo Li Xiao-Guang Qi
Northwest University, China
Wen Wang
Northwestern Polytechnical University, China
Travis Robbins
Northern New Mexico College, USA
Snorre B. Hagen
Norwegian Institute for Bioeconomy Research, Norway
Per Wegge
Norwegian University of Life Sciences, Norway
Daisuke Kondoh
Obihiro University of Agriculture, Japan
Tatsuhiko Goto Tatsuo Oshida
Obihiro University of Agriculture and Veterinary Medicine, Japan
Tomas Grim
Palacky University, Czech Republic
Sheng Li Meng Yao
Peking University, China
Wei-Hua Yue
Peking University Sixth Hospital, China
Arvin Diesmos
Philippine National Museum of Natural History, Philippines
Muhammad Sajid Nadeem
PMAS-Arid Agriculture University, Pakistan
Dmitry Bulgin
Polyclinic ME-DENT, Croatia
Gandhi Radis-Baptista
Pompeu Fabra University, Spain
Seyyed Saeed Hosseinian Yousefkhani
Razi University, Iran
Peter Jager
Research Institute Senckenberg, Germany
Felix N. Toka
Ross University School of Veterinary Medicine, USA
Cédric D'Udekem D'Acoz
Royal Belgian Institute of Natural Sciences, Belgium
Mohammad Hosein Nasr Esfahani
Royan Institute for Biotechnology, Iran
Robert Murphy
Royal Ontario Museum, Canada
Majid Eslami
Semnan University of Medical Sciences, Iran
Yu-Chun Li
Shandong University, China
Yi-Feng Gu Jin-Liang Zhao
Shanghai Ocean University, China
Yan-Feng Tong
Shenyang Normal University, China
Chihiro Takahata
Shinshu University, Japan
Hamid R. Esmaeili
Shiraz University, Iran
Qing-Yong Ni
Sichuan Agricultural University, China
Shun-De Chen Mian Hou
Sichuan Normal University, China
Zhen-Xin Fan Jing Li Zhao-Bin Song Yong-Jie Wu
Sichuan University, China
Koustubh Sharma
Snow Leopard Trust, USA
Gernot Vogel
Society for Southeast Asian Herpetology, Germany
Yong-Cheng Long
Society of Entrepreneurs & Ecology, China
Zhi-Yong Yuan
Southwest Forestry University, China
Li-Li Zhou
Southern Medical University, China
De-Shou Wang
Southwest University, China
Ji-Fan Hu
Stanford University, USA
Aletta E. Bester van der Merwe
Stellenbosch University, South Africa
Katsuhiro Konno
Stevens Institute of Technology, USA
Zhi-Tong Lyu
Sun Yat-Sen University, China
Chun-Yu Liu
SUNY Upstate Medical University, USA
John Manning
Swansea University, UK
Orjan Johansson Getinet Mekuriaw Tarekegn
Swedish University of Agricultural Sciences, Sweden
Ming Chen
Taiwan Changhua Christian Hospital, China
Wen-San Huang
Taiwan National Museum of Natural Science, China
Si-Min Lin
Taiwan Normal University, China
Wai-Yee Chan
The Chinese University of Hong Kong, China
Neal Woodman
The College of New Jersey, USA
Awal Riyanto
The Indonesian Institute of Sciences, Indonesia
Suresh Balakrishnan
The M.S. University of Baroda, India
Mike Bates
The National Museum, South Africa
Ralf Britz David J. Gower Paula Jenkins
The Natural History Museum, UK
Kichoon Lee
The Ohio State University, USA
Milton Tan
The University of Illinois at Urbana, USA
Perry L. Wood, Jr.
The University of Kansas, USA
Shiwanthi L. Ranasinghe
The University of Queensland, Australia
Christopher S. Navara
The University of Texas at San Antonio, USA
Tetsuya Shiuchi
Tokushima Universtiy, Japan
Laxman Khanal
Tribhuvan University, Nepal
Alfred Trnka
Trnava University in Trnava, Slovakia
Lin-Qi Zhang
Tsinghua University, China
Gong Chen
Tsinghua University School of Medicine, China
David Veliz
Universidad de Chile, Chile
Cristian E. Hernandez
Universidad de Concepcian, Chile
Francisco Ruiz-Raya
Universidad de Granada, Spain
Jesús Molinari
Universidad de Los Andes, Venezuela
Enrique Lanuza
Universidad de Valencia, Spain
Liliana Marcela Henao Munoz
Universidad del Tolima, Colombia
Uri Omar García-Vázquez Leticia M. Ochoa-Ochoa Hibraim A. Pérez-Mendoza
Universidad Nacional Autónoma de México, Mexico
Anibal Helios Diaz de la Vega-Perez
Universidad Autónoma de Tlaxcala, Mexico
Gabriela Cardozo
Universidad Nacional de Córdoba, Argentina
Jorgelina M. Boretto
Universidad Nacional del Comahue, Argentina
Daniel Casali
Universidade Federal de Minas Gerais, Brazil
Silvana Allodi Marcelo Britto Britto Breno Hamdan
Universidade Federal do Rio de Janeiro, Brazil
Lucas Tirloni
Universidade Federal do Rio Grande do Sul, Brazil
Cristina Cozzi
Università degli Studi di Milano, Italy
Emiliano Mori
Università degli Studi di Siena Facoltà di Scienze Matematiche Fisiche e Naturali, Italy
Giovambattista De Sarro
Universita degli Studi Magna Graecia di Catanzaro, Italy
Fabio Marroni
Università di Udine, Italy
Jorge Ramon Lopez-Olvera
Universitat Autònoma de Barcelona, Spain
Tesla A. Monson
Universitat Zürich, Switzerland
Patrick
Université de Franche-Comté, France
Hugues Fausther Bovendo
Universite Laval Faculte de Medecine, Canada
Bernard Dutrillaux
Université Paris, France
Bun Poh Keong
Universiti Pendidikan SultanIdris, Malaysia
Ina Salwany Md. Yasin Christina S. Y. Yong
Universiti Putra Malaysia, Malaysia
Muin Ma
Universiti Sains Malaysia, Malaysia
Carol Berman
University at Buffalo College of Arts and Sciences, USA
David Armisén
University Claude Bernard Lyon 1, France
Alan Myers
University College Cork, Ireland
Steve Peigneur
University Leuven (KU Leuven), Belgium
Nick Pilfold James L. Stafford
University of Alberta, Canada
Panayiotis Pafilis
University of Athens, Greece
Bettina Mahler
University of Buenos Aires, Argentina
Hong-Dian Yang
University of California, USA
Amir Ardeshir Julie B. Linden
University of California Davis, USA
Eric Pearlman
University of California, Irvine, USA
Joseph Demer
University of California, Los Angeles, USA
Andrea Strakova
University of Cambridge, UK
Ricardo Insausti
University of Castilla-La Mancha, Spain
Xue-Cong Liu Xue-Mei LV
University of Chinese Academy of Sciences, China
Joshua B. Gross
University of Cincinnati, USA
Elizabeth Herder
University of Connecticut, USA
Antonin Crumiere
University of Copenhagen Department of Biology, Denmark
Vincenzo Landi
University of Cordoba, Spain
Ioannis Koliarakis
University of Crete, Greece
Zhi-Qiang Zeng
University of Edinburgh, UK
James E. Barasa Linnet Gohole
University of Eldoret, Kenya
Valerio Donfrancesco
University of Exeter, UK
Gregor Rainer
University of Fribourg, Switzerland
Madhur Mangalam
University of Georgia, USA
Daniel Escoriza
University of Girona, Spain
Francisco Ruiz-Raya
University of Granada, Spain
Eva Edilia Avila
University of Guanajuato, Mexico
Anthony Barley David W. Greenfield
University of Hawai'i at Mānoa, USA
Yik-Hei Sung
University of Hong Kong, China
Chun-Yu Liu
University of Illinois at Chicago, USA
Perry L. Wood
University of Kansas, USA
Choo Hock Tan
University of Malaya, Malaysia
Li Ma
University of Maryland, USA
Thibault Renoir
University of Melbourne, Australia
Yu Lin
University of Michigan Medical Center, USA
Danielle H. Drabeck
University of Minnesota, USA
Wu Zhou
University of Mississippi Medical Center, USA
Miguel L. Batista Jr.
University of Mogi das Cruzes, Brazil
Domenico Fulgione
University of Naples Federico II, Italy
Kai Wang
University of Oklahoma, USA
Victor Benno Meyer-Rochow
University of Oulu, Finland
Daniela Campobello
University of Palermo, Italy
Anders Moller
University of Paris-South, France
Nigel Fraser
University of Pennsylvania, USA
Lasagna Emiliano
University of Perugia, Italy
Ci-Rong Liu Charles A. Scanga
University of Pittsburgh, USA
Thomas H.J. Burne
University of Queensland, Australia
Jacques Robert
University of Rochester, USA
Laetitia Marechal
University of Roehampton, UK
Zaida Ortega
University of Salamanca, Spain
Tiago do Prado Paim
University of Sao Paulo, Brazil
Alison Gainsbury
University of South Florida St Petersburg, USA
David C. Lee
University of South Wales, UK
Matt Neale
University of Sussex, UK
Ros Gloag
University of Sydney, Australia
Javad Javanbakht
University of Tehran, Iran
Eric N. Smith
University of Texas Arlington College of Science, USA
Christopher S. Navara
University of Texas at San Antonio, USA
Kasha Strickland
University of the Sunshine Coast, Australia
David Irwin
University of Toronto, Canada
John S. Taylor
University of Victoria, Canada
Randall C. Kyes
University of Washington, USA
Cyril C. Grueter
University of Western Australia, Australia
Charlotte Canteloup
University of Zürich, Switzerland
Christina Yong Seok Yien
University Putra Malaysia, Malaysia
Finn Hallbook Josef D. Jarhult
Uppsala University, Sweden
Hinrich Kaiser
Victor Valley College, USA
John S. Taylor
University Victoria, Canada
Christina Marie Rochus Geert Wiegertjes
Wageningen University, The Netherlands
Ryan Thummel
Wayne State University School of Medicine, USA
Taiki Ito
Wetlands International Japan, Japan
Ved Prakash Kumar Virendra Prasad Uniyal
Wildlife Institute of India, India
Fu-Qiang Xu
Wuhan Institute of Physics and Mathematics, Chinese Academy of Sciences, China
Yong-Quan Su Yi-Quan Wang
Xiamen University, China
Peng Guo
Yibin University, China
Devrim Saripinar Aksu
Yuzuncu Yil University, Turkey
Shan-Yuan Chen Jian-Jun Gao Yan Li Shu-Qun Liu Li Yu
Yunnan University, China
Tian-Xiang Gao
Zhejiang Ocean University, China
Jian-Zhong Shao
Zhejiang University, China
Ankita Rajpoot
Zoological Survey of India, India
Benjamin Tapley
ZSL London Zoo, UK
