1 Kyoto University Museum, Kyoto University, Kyoto 606-8501, Japan2 Key Laboratory of Conservation and Application in Biodiversity of South China, School of Life Sciences, Guangzhou University, Guangzhou Guangdong 510006, China3 Laboratory Animal Center, Graduate School of Medicine, Osaka City University, Osaka 545-8585, Japan4 Wildlife Research Center, Kyoto University, Sakyo, Kyoto 606-8203, Japan5 Japan Monkey Centre, Inuyama, Aichi 484-0081, Japan6 Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming Yunnan 650223, China7 Marine College, Shandong University (Weihai), Weihai Shandong 264209, China